{
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    "curator": "none",
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          "ADP",
          "MG"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
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        "salt_mM": null,
        "temperature_c": -173.15
      },
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      },
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      "reader": null,
      "source_kind": "deposit",
      "technique": "xray",
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    {
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        "buffer": null,
        "environment": "in_vitro",
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          "GDP",
          "GTP",
          "MG",
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        ],
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        "ph": null,
        "salt_mM": null,
        "temperature_c": null
      },
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        "description": "Kinesin heavy chain (chains K)",
        "labels": []
      },
      "data_format": "mmCIF + EMDB map",
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    {
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          "GTP"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
        "ph": null,
        "salt_mM": null,
        "temperature_c": null
      },
      "construct": {
        "description": "Kinesin-1 heavy chain (chains K)",
        "labels": []
      },
      "data_format": "mmCIF + EMDB map",
      "data_level": "processed",
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          "GTP",
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        ],
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        "temperature_c": null
      },
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        "description": "Kinesin-1 heavy chain (chains K)",
        "labels": []
      },
      "data_format": "mmCIF + EMDB map",
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      "source_kind": "deposit",
      "technique": "cryoem",
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    {
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          "GOL",
          "GTP",
          "MES",
          "MG",
          "SO4"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
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        "salt_mM": null,
        "temperature_c": -173.15
      },
      "construct": {
        "description": "Kinesin-1 heavy chain (chains K)",
        "labels": []
      },
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      "data_link": "https://www.rcsb.org/structure/4LNU",
      "evidence_id": "EV-PDB-4LNU-4",
      "mappings": [],
      "molecule_id": "P33176",
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        "unit": "Å",
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      },
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        "url": "https://www.rcsb.org/structure/4LNU"
      },
      "reader": null,
      "source_kind": "deposit",
      "technique": "xray",
      "title": "Nucleotide-free kinesin motor domain in complex with tubulin and a DARPin"
    },
    {
      "conditions": {
        "applied_force_pn": null,
        "buffer": null,
        "environment": "in_vitro",
        "ions": [],
        "ligands": [
          "ACT",
          "SO4",
          "SR"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
        "ph": null,
        "salt_mM": null,
        "temperature_c": -173.15
      },
      "construct": {
        "description": "Kinesin-like protein (chains A)",
        "labels": []
      },
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      "data_link": "https://www.rcsb.org/structure/5LT0",
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      "mappings": [],
      "molecule_id": "P33176",
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        "type": "resolution",
        "uncertainty": null,
        "unit": "Å",
        "value": 2.0
      },
      "provenance": {
        "deposit_id": "5LT0",
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      },
      "reader": null,
      "source_kind": "deposit",
      "technique": "xray",
      "title": "nucleotide-free kinesin-1 motor domain, P212121 crystal form"
    },
    {
      "conditions": {
        "applied_force_pn": null,
        "buffer": null,
        "environment": "in_vitro",
        "ions": [],
        "ligands": [
          "ACT",
          "SO4",
          "SR"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
        "ph": null,
        "salt_mM": null,
        "temperature_c": -173.15
      },
      "construct": {
        "description": "Kinesin-like protein (chains A,B)",
        "labels": []
      },
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        "uncertainty": null,
        "unit": "Å",
        "value": 1.949
      },
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        "doi": "10.1038/srep42558",
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        "url": "https://www.rcsb.org/structure/5LT1"
      },
      "reader": null,
      "source_kind": "deposit",
      "technique": "xray",
      "title": "nucleotide-free kinesin-1 motor domain T92V mutant, P21 crystal form"
    },
    {
      "conditions": {
        "applied_force_pn": null,
        "buffer": null,
        "environment": "in_vitro",
        "ions": [],
        "ligands": [
          "SO4"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
        "ph": 6.5,
        "salt_mM": null,
        "temperature_c": -173.15
      },
      "construct": {
        "description": "Kinesin-like protein (chains A,B,C,D,E,K)",
        "labels": []
      },
      "data_format": "mmCIF + structure factors",
      "data_level": "processed",
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      "evidence_id": "EV-PDB-5LT2-1",
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      },
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        "source": "RCSB PDB",
        "url": "https://www.rcsb.org/structure/5LT2"
      },
      "reader": null,
      "source_kind": "deposit",
      "technique": "xray",
      "title": "nucleotide-free kinesin-1 motor domain, P1 crystal form"
    },
    {
      "conditions": {
        "applied_force_pn": null,
        "buffer": null,
        "environment": "in_vitro",
        "ions": [],
        "ligands": [
          "SO4"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
        "ph": 6.5,
        "salt_mM": null,
        "temperature_c": -173.15
      },
      "construct": {
        "description": "Kinesin-1 heavy chain (chains A,B,C,D,E,K)",
        "labels": []
      },
      "data_format": "mmCIF + structure factors",
      "data_level": "processed",
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      "evidence_id": "EV-PDB-5LT3-1",
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        "unit": "Å",
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      },
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        "doi": "10.1038/srep42558",
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        "source": "RCSB PDB",
        "url": "https://www.rcsb.org/structure/5LT3"
      },
      "reader": null,
      "source_kind": "deposit",
      "technique": "xray",
      "title": "nucleotide-free kinesin-1 motor domain T87A mutant, P1 crystal form"
    },
    {
      "conditions": {
        "applied_force_pn": null,
        "buffer": null,
        "environment": "in_vitro",
        "ions": [],
        "ligands": [
          "SO4"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
        "ph": 6.5,
        "salt_mM": null,
        "temperature_c": -173.15
      },
      "construct": {
        "description": "Kinesin-1 heavy chain (chains A,B,C,D,E,K)",
        "labels": []
      },
      "data_format": "mmCIF + structure factors",
      "data_level": "processed",
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      "evidence_id": "EV-PDB-5LT4-1",
      "mappings": [],
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      },
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        "url": "https://www.rcsb.org/structure/5LT4"
      },
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      "source_kind": "deposit",
      "technique": "xray",
      "title": "nucleotide-free kinesin-1 motor domain T92V mutant, P1 crystal form"
    },
    {
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        "buffer": null,
        "environment": "in_vitro",
        "ions": [],
        "ligands": [
          "GDP",
          "GTP",
          "MG",
          "TA1"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
        "ph": null,
        "salt_mM": null,
        "temperature_c": null
      },
      "construct": {
        "description": "Kinesin-like protein,Kinesin-1 heavy chain (chains K)",
        "labels": []
      },
      "data_format": "mmCIF + EMDB map",
      "data_level": "processed",
      "data_link": "https://www.rcsb.org/structure/7TR2",
      "evidence_id": "EV-PDB-7TR2-1",
      "mappings": [],
      "molecule_id": "P33176",
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      },
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        "doi": "10.1038/s41467-022-31794-3",
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        "source": "RCSB PDB",
        "url": "https://www.rcsb.org/structure/7TR2"
      },
      "reader": null,
      "source_kind": "deposit",
      "technique": "cryoem",
      "title": "Apo CaKip3[2-436]-L2-mutant(HsKHC) in complex with a microtubule"
    },
    {
      "conditions": {
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        "buffer": null,
        "environment": "in_vitro",
        "ions": [],
        "ligands": [
          "ATP",
          "G2P",
          "GTP"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
        "ph": null,
        "salt_mM": null,
        "temperature_c": null
      },
      "construct": {
        "description": "Kinesin-1 heavy chain (chains S,T,U,V,W,X,Y,Z,a)",
        "labels": []
      },
      "data_format": "mmCIF + EMDB map",
      "data_level": "processed",
      "data_link": "https://www.rcsb.org/structure/8IXA",
      "evidence_id": "EV-PDB-8IXA-3",
      "mappings": [],
      "molecule_id": "P33176",
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        "type": "resolution",
        "uncertainty": null,
        "unit": "Å",
        "value": 4.2
      },
      "provenance": {
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        "doi": "10.3724/abbs.2023130",
        "licence": "CC0 1.0 (wwPDB archive data)",
        "retrieved": "2026-10-19",
        "source": "RCSB PDB",
        "url": "https://www.rcsb.org/structure/8IXA"
      },
      "reader": null,
      "source_kind": "deposit",
      "technique": "cryoem",
      "title": "GMPCPP-Alpha1A/Beta2A-microtubule decorated with kinesin non-seam region"
    },
    {
      "conditions": {
        "applied_force_pn": null,
        "buffer": null,
        "environment": "in_vitro",
        "ions": [],
        "ligands": [
          "ATP",
          "G2P",
          "GTP"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
        "ph": null,
        "salt_mM": null,
        "temperature_c": null
      },
      "construct": {
        "description": "Kinesin-1 heavy chain (chains g,k,l,m)",
        "labels": []
      },
      "data_format": "mmCIF + EMDB map",
      "data_level": "processed",
      "data_link": "https://www.rcsb.org/structure/8IXB",
      "evidence_id": "EV-PDB-8IXB-3",
      "mappings": [],
      "molecule_id": "P33176",
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        "unit": "Å",
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      },
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        "doi": "10.3724/abbs.2023130",
        "licence": "CC0 1.0 (wwPDB archive data)",
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        "source": "RCSB PDB",
        "url": "https://www.rcsb.org/structure/8IXB"
      },
      "reader": null,
      "source_kind": "deposit",
      "technique": "cryoem",
      "title": "GMPCPP-Alpha1A/Beta2A-microtubule decorated with kinesin seam region"
    },
    {
      "conditions": {
        "applied_force_pn": null,
        "buffer": null,
        "environment": "in_vitro",
        "ions": [],
        "ligands": [
          "ATP",
          "G2P",
          "GTP"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
        "ph": null,
        "salt_mM": null,
        "temperature_c": null
      },
      "construct": {
        "description": "Kinesin-1 heavy chain (chains h,l,p,q)",
        "labels": []
      },
      "data_format": "mmCIF + EMDB map",
      "data_level": "processed",
      "data_link": "https://www.rcsb.org/structure/8IXG",
      "evidence_id": "EV-PDB-8IXG-3",
      "mappings": [],
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        "unit": "Å",
        "value": 4.4
      },
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        "doi": "10.3724/abbs.2023130",
        "licence": "CC0 1.0 (wwPDB archive data)",
        "retrieved": "2026-10-19",
        "source": "RCSB PDB",
        "url": "https://www.rcsb.org/structure/8IXG"
      },
      "reader": null,
      "source_kind": "deposit",
      "technique": "cryoem",
      "title": "GMPCPP-Alpha4A/Beta2A-microtubule decorated with kinesin seam region"
    },
    {
      "conditions": {
        "applied_force_pn": null,
        "buffer": null,
        "environment": "in_vitro",
        "ions": [],
        "ligands": [
          "ANP",
          "GDP",
          "GTP",
          "MG",
          "TA1"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
        "ph": null,
        "salt_mM": null,
        "temperature_c": null
      },
      "construct": {
        "description": "Kif5b Tail (chains t)",
        "labels": []
      },
      "data_format": "mmCIF + EMDB map",
      "data_level": "processed",
      "data_link": "https://www.rcsb.org/structure/8RHB",
      "evidence_id": "EV-PDB-8RHB-4",
      "mappings": [],
      "molecule_id": "P33176",
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        "uncertainty": null,
        "unit": "Å",
        "value": 3.0
      },
      "provenance": {
        "deposit_id": "8RHB",
        "doi": "10.1038/s41467-025-61498-3",
        "licence": "CC0 1.0 (wwPDB archive data)",
        "retrieved": "2026-10-19",
        "source": "RCSB PDB",
        "url": "https://www.rcsb.org/structure/8RHB"
      },
      "reader": null,
      "source_kind": "deposit",
      "technique": "cryoem",
      "title": "Microtubule-associated kinesin-1 tail complex bound to AMPPNP, single-headed state"
    },
    {
      "conditions": {
        "applied_force_pn": null,
        "buffer": null,
        "environment": "in_vitro",
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        "ligands": [
          "ANP",
          "GDP",
          "GTP",
          "MG",
          "TA1"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
        "ph": null,
        "salt_mM": null,
        "temperature_c": null
      },
      "construct": {
        "description": "Kinesin-1 heavy chain (chains K,T)",
        "labels": []
      },
      "data_format": "mmCIF + EMDB map",
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        "description": "Kinesin-1 heavy chain (chains K)",
        "labels": []
      },
      "data_format": "mmCIF + structure factors",
      "data_level": "processed",
      "data_link": "https://www.rcsb.org/structure/4HNA",
      "evidence_id": "EV-PDB-4HNA-4",
      "mappings": [],
      "molecule_id": "P33176",
      "observable": {
        "type": "resolution",
        "uncertainty": null,
        "unit": "Å",
        "value": 3.19
      },
      "provenance": {
        "deposit_id": "4HNA",
        "doi": "10.1038/nsmb.2624",
        "licence": "CC0 1.0 (wwPDB archive data)",
        "retrieved": "2026-10-19",
        "source": "RCSB PDB",
        "url": "https://www.rcsb.org/structure/4HNA"
      },
      "reader": null,
      "source_kind": "deposit",
      "technique": "xray",
      "title": "Kinesin motor domain in the ADP-MG-ALFX state in complex with tubulin and a DARPIN"
    },
    {
      "conditions": {
        "applied_force_pn": null,
        "buffer": null,
        "environment": "in_vitro",
        "ions": [],
        "ligands": [
          "ADP",
          "MG",
          "SO4"
        ],
        "notes": "Entry-level ligand list; oligosaccharides appear as branched entities. pH and temperature are null when not deposited in machine-readable fields.",
        "ph": 7.5,
        "salt_mM": null,
        "temperature_c": -173.15
      },
      "construct": {
        "description": "KINESIN HEAVY CHAIN (chains A)",
        "labels": []
      },
      "data_format": "mmCIF + structure factors",
      "data_level": "processed",
      "data_link": "https://www.rcsb.org/structure/1MKJ",
      "evidence_id": "EV-PDB-1MKJ-1",
      "mappings": [],
      "molecule_id": "P33176",
      "observable": {
        "type": "resolution",
        "uncertainty": null,
        "unit": "Å",
        "value": 2.7
      },
      "provenance": {
        "deposit_id": "1MKJ",
        "doi": null,
        "licence": "CC0 1.0 (wwPDB archive data)",
        "retrieved": "2026-10-19",
        "source": "RCSB PDB",
        "url": "https://www.rcsb.org/structure/1MKJ"
      },
      "reader": null,
      "source_kind": "deposit",
      "technique": "xray",
      "title": "Human Kinesin Motor Domain With Docked Neck Linker"
    },
    {
      "conditions": {
        "applied_force_pn": null,
        "buffer": null,
        "environment": "in_vitro",
        "ions": [],
        "ligands": [],
        "notes": "From the first sample condition list of the entry.",
        "ph": 7.4,
        "salt_mM": null,
        "temperature_c": 9.85
      },
      "construct": {
        "description": "BMRB data: 13C chemical shifts, 15N chemical shifts",
        "labels": []
      },
      "data_format": "NMR-STAR",
      "data_level": "processed",
      "data_link": "https://bmrb.io/data_library/summary/index.php?bmrbId=30936",
      "evidence_id": "EV-BMRB-30936",
      "mappings": [],
      "molecule_id": "P33176",
      "observable": {
        "type": "chemical_shift",
        "uncertainty": null,
        "unit": null,
        "value": null
      },
      "provenance": {
        "deposit_id": "bmr30936",
        "doi": "10.13018/BMR30936",
        "licence": "CC0 1.0 (verify)",
        "retrieved": "2026-10-19",
        "source": "BMRB",
        "url": "https://bmrb.io/data_library/summary/index.php?bmrbId=30936"
      },
      "reader": null,
      "source_kind": "deposit",
      "technique": "nmr_shifts",
      "title": "Atomic-Resolution Structure of Kinesin-1 Motor Domain in Complex with Polymeric Microtubules by Magic Angle Spinning NMR"
    },
    {
      "conditions": {
        "applied_force_pn": null,
        "buffer": null,
        "environment": "in_silico",
        "ions": [],
        "ligands": [],
        "notes": "Computed prediction; no experimental conditions.",
        "ph": null,
        "salt_mM": null,
        "temperature_c": null
      },
      "construct": {
        "description": "AlphaFold Monomer v2.0 pipeline, residues 1-963",
        "labels": []
      },
      "data_format": "mmCIF, PAE JSON",
      "data_level": "processed",
      "data_link": "https://alphafold.ebi.ac.uk/entry/P33176",
      "evidence_id": "EV-AFDB-AF-P33176-F1",
      "mappings": [],
      "molecule_id": "P33176",
      "observable": {
        "type": "plddt",
        "uncertainty": null,
        "unit": "pLDDT",
        "value": 78.56
      },
      "provenance": {
        "deposit_id": "AF-P33176-F1",
        "doi": null,
        "licence": "CC BY 4.0",
        "retrieved": "2026-10-19",
        "source": "AlphaFold DB",
        "url": "https://alphafold.ebi.ac.uk/entry/P33176"
      },
      "reader": null,
      "source_kind": "computed",
      "technique": "af_prediction",
      "title": "AlphaFold DB prediction AF-P33176-F1"
    },
    {
      "conditions": {
        "applied_force_pn": null,
        "buffer": null,
        "environment": "in_silico",
        "ions": [],
        "ligands": [],
        "notes": "Simulation; temperature is the thermostat setting.",
        "ph": null,
        "salt_mM": null,
        "temperature_c": 26.85
      },
      "construct": {
        "description": "start 1BG2; Amber Parm99",
        "labels": []
      },
      "data_format": "trajectory",
      "data_level": "processed",
      "data_link": "https://mmb.mddbr.eu/#/id/A006A/overview",
      "evidence_id": "EV-MDDB-A006A",
      "mappings": [],
      "molecule_id": "P33176",
      "observable": {
        "type": "trajectory",
        "uncertainty": null,
        "unit": "ns",
        "value": 10.0
      },
      "provenance": {
        "deposit_id": "A006A",
        "doi": "10.1016/j.str.2010.07.013",
        "licence": "This trajectory dataset is released under a Creative Commons Attribution 4.0 International Public License",
        "retrieved": "2026-10-19",
        "source": "MDDB",
        "url": "https://mmb.mddbr.eu/#/id/A006A/overview"
      },
      "reader": null,
      "source_kind": "computed",
      "technique": "md",
      "title": "MoDEL 1BG2"
    }
  ],
  "models": [],
  "molecule": {
    "components": [],
    "domains": [
      {
        "end": 325,
        "name": "Kinesin motor",
        "start": 8
      }
    ],
    "gene": "KIF5B",
    "identifiers": {
      "genbank": null,
      "isoform": null,
      "nakb": null,
      "pdb": [],
      "rfam": null,
      "rnacentral": null,
      "uniprot": "P33176"
    },
    "ligands": [
      "ATP"
    ],
    "modifications": [
      {
        "kind": "ptm",
        "name": "N-acetylalanine",
        "position": 2,
        "source": "UniProtKB"
      },
      {
        "kind": "ptm",
        "name": "Phosphoserine",
        "position": 933,
        "source": "UniProtKB"
      },
      {
        "kind": "ptm",
        "name": "Omega-N-methylarginine",
        "position": 956,
        "source": "UniProtKB"
      }
    ],
    "molecule_id": "P33176",
    "molecule_type": "protein",
    "name": "Kinesin-1 heavy chain",
    "organism": {
      "name": "Homo sapiens",
      "taxon_id": 9606
    },
    "provenance": {
      "deposit_id": "P33176",
      "doi": null,
      "licence": "CC BY 4.0 (UniProt)",
      "retrieved": "2026-10-19",
      "source": "UniProtKB",
      "url": "https://www.uniprot.org/uniprotkb/P33176/entry"
    },
    "sequence": "MADLAECNIKVMCRFRPLNESEVNRGDKYIAKFQGEDTVVIASKPYAFDRVFQSSTSQEQVYNDCAKKIVKDVLEGYNGTIFAYGQTSSGKTHTMEGKLHDPEGMGIIPRIVQDIFNYIYSMDENLEFHIKVSYFEIYLDKIRDLLDVSKTNLSVHEDKNRVPYVKGCTERFVCSPDEVMDTIDEGKSNRHVAVTNMNEHSSRSHSIFLINVKQENTQTEQKLSGKLYLVDLAGSEKVSKTGAEGAVLDEAKNINKSLSALGNVISALAEGSTYVPYRDSKMTRILQDSLGGNCRTTIVICCSPSSYNESETKSTLLFGQRAKTIKNTVCVNVELTAEQWKKKYEKEKEKNKILRNTIQWLENELNRWRNGETVPIDEQFDKEKANLEAFTVDKDITLTNDKPATAIGVIGNFTDAERRKCEEEIAKLYKQLDDKDEEINQQSQLVEKLKTQMLDQEELLASTRRDQDNMQAELNRLQAENDASKEEVKEVLQALEELAVNYDQKSQEVEDKTKEYELLSDELNQKSATLASIDAELQKLKEMTNHQKKRAAEMMASLLKDLAEIGIAVGNNDVKQPEGTGMIDEEFTVARLYISKMKSEVKTMVKRCKQLESTQTESNKKMEENEKELAACQLRISQHEAKIKSLTEYLQNVEQKKRQLEESVDALSEELVQLRAQEKVHEMEKEHLNKVQTANEVKQAVEQQIQSHRETHQKQISSLRDEVEAKAKLITDLQDQNQKMMLEQERLRVEHEKLKATDQEKSRKLHELTVMQDRREQARQDLKGLEETVAKELQTLHNLRKLFVQDLATRVKKSAEIDSDDTGGSAAQKQKISFLENNLEQLTKVHKQLVRDNADLRCELPKLEKRLRATAERVKALESALKEAKENASRDRKRYQQEVDRIKEAVRSKNMARRGHSAQIAKPIRPGQHPAASPTHPSAIRGGGAFVQNSQPVAVRGGGGKQV",
    "sequence_length": 963
  },
  "molecule_id": "P33176",
  "states": [],
  "transitions": []
}